1. Prichard A, Cook PF, Spivak M, Chhibber R, Berns GS: Awake fMRI reveals brain regions for novel word detection in dogs. Frontiers in Neuroscience 12:737, 2018. PDFLink.
  2. Prichard A, Chhibber R, Athanassiades K, Spivak M, Berns GS: Fast neural learning in dogs: a multimodal fMRI study. Sci Rep 8:14614, 2018. PDF. Link.
  3. Berns GS, Spivak M, Nemanic S, Northrup N: Clinical findings in dogs trained for awake-MRI. Frontiers in Veterinary Science 5:209, 2018. PDFLink
  4. Cook P, Prichard A, Spivak M, Berns GS: Jealousy in dogs? Evidence from brain imaging. Animal Sentience 22:1, 2018. PDFLink.
  5. Cook PF, Berns GS, Colegrove K, Johnson S, Gulland F: Post-mortem DTI reveals altered hippocampal connectivity in wild sea lions diagnosed with chronic toxicosis from algal exposure. J Comp Neurol 526:216-228, 2018. Online Sept 2017.
  6. Engelmann JB, Berns GS, Dunlop BW: Hyper-responsivity to losses in the anterior insula during economic choice scales with depression severity. Psychol Med Jun 7:1-13, 2017.
  7. Berns GS, Brooks AM, Spivak M, Levy K: Functional MRI in awake dogs predicts suitability for assistance work. Sci Rep 7:43704, 2017. PDFLink.
  8. Berns GS & Ashwell KWS: Reconstruction of the cortical maps of the Tasmanian tiger and comparison to the Tasmanian devil. PLoS ONE 12(1):e0168993, 2017. PDFLink.
  9. Cook PF, Prichard A, Spivak M, Berns GS: Awake canine fMRI predicts dogs' preference for praise versus food. Soc Cog Affect Neurosci, 11:1853-1862, 2016. PDFLink.
  10. Cook PF, Spivak M, Berns G: Neurobehavioral evidence for individual differences in canine cognitive control: an awake fMRI study. Anim Cogn, 19:867-878, 2016. LinkPDF.
  11. Dilks DD, Cook P, Weiller SK, Berns HP, Spivak M, Berns GS: Awake fMRI reveals a specialized region in dog temporal cortex for face processing. PeerJ, 3:e1115, 2015. LinkPDF.
  12. Berns GS, Cook PF, Foxley S, Jbabdi S, Miller KL, Marino L: Diffusion tensor imaging of dolphin brains reveals direct auditory pathway to temporal lobe. Proc. R. Soc. B 282:20151203, 2015. Link.
  13. Cook PF, Spivak M, Berns GS: One pair of hands is not like another: caudate BOLD response in dogs depends on signal source and canine temperament. PeerJ, 2:e596, 2014. PDFLink.
  14. Pincus M, LaViers L, Prietula MJ, Berns G: The conforming brain and deontological resolve. PLoS ONE, 9(8):e106061, 2014. Link.
  15. Ekins WG, Brooks AM, Berns GS: The neural correlates of contractual risk and penalty framing. J. Risk & Uncertainty, 49:125-140, 2014. PDF.
  16. Barton J, Berns GS, Brooks AM: The neuroscience behind the stock market's reaction to corporate earnings news. The Accounting Review, 89:1945-1977, 2014.
  17. Berns GS, Brooks AM, Spivak M: Scent of the familiar: an fMRI study of canine brain responses to familiar and unfamiliar human and dog odors. Behavioural Processes, 2014. Link.
  18. Berns GS, Blaine K, Prietula MJ, Pye BE: Short and long term effects of a novel on connectivity in the brain. Brain Connectivity 3:590-600, 2013. Link. Most downloaded paper in the journal in 2017.
  19. Ekins WG, Caceda R, Capra CM, Berns GS: You can't gamble on others: dissociable systems for strategic uncertainty and risk in the brain. J. Econ. Behav. Org.94:222-233, 2013. PDF.
  20. Berns GS, Brooks A, Spivak M: Replicability and heterogeneity of awake unrestrained canine fMRI responses. PLoS ONE 8(12):e81698, 2013. Link.
  21. Capra CM, Jiang B, Engelmann JB, Berns GS: Can personality explain heterogeneity in probability distortions? J. Neurosci. Psychol. Econ. 6:151-166, 2013. PDF.
  22. Berns GS, Brooks AM, Spivak M: Functional MRI in awake unrestrained dogs. PLoS ONE 7(5):e38027, 2012. LinkPDF.
  23. Engelmann JB, Moore S, Capra CM, Berns GS: Differential neurobiological effects of expert advice on risky choice in adolescents and adults. Soc Cogn Affect Neurosci, 7:557-567, 2012. PDF.
  24. Berns GS, Bell E, Capra CM, Prietula MJ, Moore S, Anderson B, Ginges J, Atran S: The price of your soul: neural evidence for the non-utilitarian representation of sacred values. Phil. Trans. Royal Soc. B, 367:754-762, 2012. PDFHTML.
  25. Brooks AM, Capra CM, Berns GS: Neural insensitivity to upticks in value is associated with the disposition effect. Neuroimage, 59:4086-4093, 2012. PDF.
  26. Berns GS and Bell E: Striatal topography of probability and magnitude information for decisions under uncertainty. Neuroimage, 59:3166-3172, 2012. PDF.
  27. Berns GS and Moore SE: A neural predictor of cultural popularity. J Consumer Psychol. 22:154-160, 2012. doi:10.1016/j.jcps.2011.05.001.
  28. Brooks AM, Pammi VSC, Noussair C, Capra CM, Engelmann JB, Berns GS: From bad to worse: striatal coding of the relative value of painful decisions. Front. Neurosci doi: 10.3389/fnins.2010.00176, 2010. PDF.
  29. Berns GS, Capra CM, Moore S, Noussair C: Neural mechanisms of the influence of popularity on adolescent ratings of music. Neuroimage 49:2687-2696, 2010. PDF.
  30. Berns GS, Moore S, Capra CM: Adolescent engagement in dangerous behaviors is associated with increased white matter maturity of frontal cortex. PLoS One 4(8):e6773, 2009. LinkPDF.
  31. Engelmann JB, Capra CM, Noussair C, Berns GS: Expert financial advice neurobiologically "offloads" financial decision-making under risk. PLoS One 4:e4957, 2009. LinkPDF.
  32. Berns GS, Capra CM, Chappelow J, Moore S, Noussair C: Nonlinear neurobiological probability weighting functions for aversive outcomes. Neuroimage 39:2047-2057, 2008 (epub Nov 2007). PDF.
  33. Chandrasekhar PVS, Capra CM, Moore S, Noussair C, Berns GS: Neurobiological regret and rejoice functions for aversive outcomes. Neuroimage 39:1472-1484 (epub Nov 2007). PDF.
  34. Berns GS, Capra CM, Moore S, Noussair C: A shocking experiment: new evidence on probability weighting and common ratio violations. Judgment & Decision Making 2:234-242, 2007. PDF.
  35. Capuron L, Pagnoni G, Demetrashvili MF, Lawson DH, Fornwalt FB, Woolwine B, Berns GS, Nemeroff CB, Miller AH: Basal ganglia hypermetabolism and symptoms of fatigue during interferon-alpha therapy. Neuropsychopharmacology 32:2394-2392 , 2007.
  36. Berns GS, Chappelow J, Cekic M, Zink CF, Pagnoni G, Martin-Skurski ME: Neurobiologic substrates of dread. Science, 312:754-758, 2006. PDFSupporting Materials.
  37. Zink CF, Pagnoni G, Chappelow JC, Martin-Skurski ME, Berns GS: Human striatal activation reflects degree of stimulus saliency. Neuroimage 29:977-983, 2006. PDF.
  38. Berns GS, Chappelow JC, Zink CF, Pagnoni G, Martin-Skurski ME, Richards R: Neurobiological correlates of social conformity and independence during mental rotation. Biol. Psychiatry 58:245-253, 2005. PDF.
  39. Capuron L, Pagnoni G, Demetrashvili M, Woolwine BJ, Nemeroff CB, Berns GS, Miller AH: Anterior cingulate activation and error processing during interferon-alpha treatment. Biol. Psychiatry 58:190-196, 2005. PDF.
  40. Zink CF, Pagnoni G, Martin-Skurski ME, Chappelow JC, Berns GS: Human striatal response to monetary reward depends on saliency. Neuron 42:509-517, 2004. PDF.
  41. Zink CF, Pagnoni G, Martin ME, Dhamala M, Berns GS: Human striatal response to salient non-rewarding stimuli. J. Neurosci. 23:8092-8097, 2003. PDFAccompanying editorial.
  42. Dhamala M, Pagnoni G, Wiesenfeld K, Zink CF, Martin M, Berns GS: Neural correlates of the complexity of rhythmic finger tapping. Neuroimage 20:918-926, 2003. PDF.
  43. McClure SM, Berns GS, Montague PR: Temporal prediction errors in a passive learning task activate human striatum. Neuron 38: 339-346, 2003. PDF.
  44. Montague PR, Berns GS, Cohen JD, McClure SM, Pagnoni G, Dhamala M, Wiest M, Karpov I, King RD, Apple N, Fisher RE: Hyperscanning: Simultaneous fMRI during linked social interactions, Neuroimage 16:1159-1164, 2002. PDF.
  45. Rilling JK, Gutman DA, Zeh TR, Pagnoni G, Berns GS, Kilts CD: A neural basis for social cooperation. Neuron, 35:395-405, 2002. PDF.
  46. Dhamala M, Pagnoni G, Wiesenfeld K, Berns GS: Measurements of brain activity complexity for varying mental loads. Phys. Rev. E., 65:041917(7), 2002. PDF.
  47. Pagnoni G, Zink CF, Montague PR, Berns GS: Activity in human ventral striatum locked to errors of reward prediction. Nat. Neurosci., 5:97-98, 2002. PDF.
  48. Berns GS, Martin M, Proper S: Limbic hyperreactivity in bipolar II disorder. Am. J. Psychiatry159:304-306, 2002. PDF.
  49. Bischoff-Grethe A, Martin M, Mao H, Berns GS: The context of uncertainty modulates the subcortical response to predictability. J. Cogn. Neurosci., 13:986-993, 2001. PDF.
  50. Berns GS, McClure SM, Pagnoni G, Montague PR: Predictability modulates human brain response to reward. J. Neurosci., 21:2793-2798, 2001. PDF.
  51. Bischoff-Grethe A, Proper SM, Mao H, Daniels KA, Berns GS: Conscious and unconscious processing of nonverbal predictability in Wernicke's area.  J. Neurosci. 20(5):1975-1981, 2000. PDF.
  52. Berns GS, Song AW, Mao H:  Continuous functional magnetic resonance imaging reveals dynamic nonlinearities in "dose-response" curves for finger opposition. J. Neurosci. 19:RC17:1-6, 1999. PDF.
  53. Nemeroff CB, Kilts CD, Berns GS:  Functional brain imaging: Twenty-first century phrenology or psychobiological advance for the millennium? Am. J. Psychiatry 156:671-673, 1999.
  54. Berns GS and Sejnowski TJ: A computational model of how the basal ganglia produce sequences. J. Cogn. Neurosci., 10:108-121, 1998. PDF.
  55. Berns GS, Cohen JD, Mintun MA: Brain regions responsive to novelty in the absence of awareness. Science, 276:1272-1275, 1997. PDF
  56. Hull ML, Berns GS, Varma H, Patterson HA: Strain in the medial collateral ligament of the human knee under single and combined loads. J. Biomech. 29:199-206, 1996.
  57. Berns GS, Dayan P, Sejnowski TJ: A correlational model for the development of disparity selectivity in visual cortex that depends on prenatal and postnatal phases. Proc. Natl. Acad. Sci. USA 90:8277-8281, 1993. PDF.
  58. Berns GS and Howell SM: Roofplasty requirements in vitro for different tibial hole placements in anterior cruciate ligament reconstructions. Am. J. Sports Med. 21:292-298, 1993.
  59. Berns GS, Howell SM, Farley TE: The accuracy of signal intensity measurements in magnetic resonance imaging as evaluated within the knee. Magn. Reson. Imag. 10:573-578, 1992.
  60. Berns GS, Hull ML, Patterson HA: Strain in the anteromedial bundle of the anterior cruciate ligament under combination loading. J. Orthop. Res. 10:167-176, 1992.
  61. Howell SM, Berns GS, Farley TE: Unimpinged and impinged anterior cruciate ligament grafts: MR signal intensity measurements. Radiology 179:639-643, 1991.
  62. Berns GS, Hull ML, Patterson HA: Implementation of a five degree of freedom automated system to determine knee flexibility in vitro. J. Biomech. Eng. 112:392-400, 1990.
  63. Berns MW, Berns GS, Coffey J, Wile AG: Exposure (dose) tables for hematoporphyrin derivative photoradiation therapy. Lasers in Surgery & Med. 4:107-131, 1984.
  64. Berns GS and Berns MW: Computer-based tracking of living cells. Exp. Cell Res. 142:103-109, 1982.
  65. Berns MW, Aist J, Edwards J, Strahs K, Girton J, McNeill P, Rattner JB, Kitzes M, Hammer-Wilson M, Liaw L-H, Siemens A, Koonce M, Peterson S, Brenner S, Burt J, Walter R, Bryant PJ, van Dyk D, Coulombe J, Cahill T, Berns GS: Laser microsurgery in cell and developmental biology. Science 213:505-513, 1981.


 Review Articles

  1. Berns GS & Cook PF: Why Did the Dog Walk into the MRI? Current Directions Psychol Sci, 25:363-369, 2016. Link.
  2. Cook PF, Brooks A, Spivak M, Berns GS: Regional brain activations in awake unrestrained dogs. J Vet Behav: Clin Appl Res, 16:104-112, 2016. Link.
  3. Brooks AM and Berns GS: Aversive stimuli and loss in the mesocortical dopamine system. Trends Cogn. Sci. 17:281-286, 2013.
  4. Yoon C, Gonzalez R, Bechara A, Berns GS, Dagher AA, Dube L, Huettel SA, Kable JW, Liberzon I, Plassmann H, Spence C: Decision neuroscience and consumer decision making. Mark Lett 23:473-485, 2012. PDF.
  5. Berns GS and Atran S: The biology of cultural conflict. Phil. Trans. Royal Soc. B, 367:633-639, 2012. PDF. HTML.
  6. Ariely D and Berns GS: Neuromarketing: The hope and hype of neuoroimaging in business. Nat. Rev. Neurosci., 11:284-292, 2010. PDF.
  7. Berns GS, Capra CM, Moore S, Noussair C: Three studies on the neuroeconomics of decision-making when payoffs are real and negative. Adv Health Econ Health Serv Res 20:1-29, 2008.
  8. Berns GS, Laibson D, Loewenstein G: Intertemporal choice -- toward an integrated framework, Trends Cogn. Sci., 11:482-488, 2007. PDF.
  9. Berns GS, Capra CM, Noussair C: Receptor theory and biological constraints on value. Ann. N.Y. Acad. Sci. 1104:301-309, 2007. PDF.
  10. Berns GS: Price, placebo, and the brain. J. Marketing Res. XLII:399-400, 2005. PDF.
  11. Berns GS: Something funny happened to reward. Trends Cogn. Sci. 8:193-194, 2004. PDF.
  12. Berns GS: Neural game theory and the search for rational agents in the brain. Behav. Brain Sci. 26:155-156, 2003.
  13. Berns GS and Nemeroff CB: The neurobiology of bipolar disorder. Am. J. Med. Genet C 123C:76-84, 2003. PDF.
  14. Montague PR and Berns GS: Neural economics and the biological substrates of valuation. Neuron 36:265-284, 2002. PDF.
  15. Mao H and Berns GS: MRI in the study of brain functions: clinical perspectives. Medicamundi 46:28-38, 2002. PDF.
  16. Berns GS and Nemeroff CB:  Extreme measures. The neurobiology of bipolar disorder.  Odyssey 5:36-41, 1999.
  17. Berns GS: Functional neuroimaging.  Life Sci 65:2531-2540, 1999. PDF.

Other Publications (Thesis and Dissertation)

  1. Berns GS: Force distribution patterns in the foot during running. A.B. Thesis, Dept. of Physics, Princeton University, 1986.
  2. Berns GS: Strain within the anterior cruciate and medial collateral ligaments of the human knee under combination loading. Ph.D. Dissertation, Biomedical Engineering, University of California, Davis, 1990.


  • 5/97 National Public Radio. Interview on learning without awareness.
  • 5/97 BBC. Interviewed on learning without awareness.
  • 7/97 Insight magazine. Quoted on subconscious learning in interview with Gavin de Becker and his book, "The Gift of Fear."
  • 9/97 Psychology Today. Interview on, "Learning but not Trying."
  • 5/99 Georgia Tech / Emory MiniMedical School lecture on "Imaging the Mind."
  • 7/99 WSB-TV Atlanta. Interviewed for special production on Depression.
  • 12/00 CNN-FYI. Live webcast for high-school Q&A on "Brain Structure and Function".
  • 4/01 New York Times, Washington Post, LA Times,,,, WebMD, Science, Scientific American, Reuters Health. Write-ups on the role of surprise in reward processing.
  • 7/01 Self Magazine. Article on surprise and reward.
  • 10/01 Prevention Magazine. Article on surprise and reward.
  • 2/02 New York Times full-length article on neural economics.
  • 3/02 Nature News & Views on Hyperscanning.
  • 7/02 New York Times full length article on neural basis of cooperation.
  • 7/02 International Herald Tribune, Front Page of National Post (Canada), Reuters, WIRED, BBC, FOXNews, Atlanta Journal-Constitution coverage of neural basis of cooperation.
  • 8/02 Chicago Tribune story on the evolution of cooperation.
  • 9/02 "Madison Avenue and Your brain"
  • 10/02 Money Magazine. Cover story on "Your Money & Your Brain"
  • 10/02 Reader's Digest.
  • 6/03 NY Times "Brain experts follow the money"
  • 9/03 Forbes "In search of the buy button"
  • 9/03 Financial Times "In search of the inside story of economics"
  • 3/04 O Magazine, article on novelty and happiness
  • 5/04 AP article on working for money
  • 5/04 CNN America Morning live interview on working for money
  • 4/05 First article on Surprise
  • 6/05 NY Times "What Other People Say May Change What You See"
  • 9/05 LA Times "For True Fulfillment, Seek Satisfaction, Not Happiness"
  • 9/05 O Magazine, Fortune, Glamour, Psychology Today, reviews of Satisfaction
  • 9/05 Town Hall Seattle Science Lecture on "Satisfaction"
  • 9/05 Microsoft Headquarters, lecture on "Satisfaction"
  • 9/05 NPR - Science Friday, "The Science of Satisfaction"
  • 12/05 Atlanta Magazine: book excerpt
  • 12/05 Wall Street Journal article on "shopping high"
  • 1/06 ABC Primetime segment on conformity
  • 2/06 Washington Post: review of Satisfaction
  • 5/06 New York Times, AP, NPR, BBC, and multiple outlets: coverage of neurobiology of dread
  • 3/07 The Space Show: interview on risky and decision-making in space exploration
  • 5/07 ReachMD (XM 233): interview on risky decision making
  • 5/07 Fox Atlanta: TV news segment on the teenage brain